HP describes these as 49.XXXX errors (without the suffix numbers after the . (dot)). The general description is that a critical firmware error has occurred, but it's usually not firmware, especially if the printer has been fine until recently.
Usually it's corruption in the print stream for a particular print job, so it could be the program (does this happen with print jobs from a particular application or from all applications; have you installed new software or upgraded it recently for the application that causes the error if it's always the same application?), the driver (have you recently upgraded it?), data corruption in the connection to the printer (bad network or USB cable, etc.), or maybe the firmware.
If they're random, and you can clear them and bring the printer back to ready by clearing the print job out of the spooler, turning off the printer, then turning it on again, it's most likely corrupt data in a print job introduced by one or more of the factors above. If they're infrequent and not too much of a bother, you can just do that.
More information - here's an HP support document on troubleshooting 49.XXXX errors:
http://h20000.www2.hp.com/bizsupport/TechSupport/D
Sometimes but rarely, they can be corrected with a firmware upgrade IF your model's firmware is flashable, and IF the problem is in the list of fixes.
Approach upgrading firmware cautiously since a firmware upgrade that doesn't install correctly can render the printer inoperable until you replace the firmware dimm with a new one.
To check for an available upgrade, go to hp.com.
Click Support & Drivers.
Select "Download drivers and software (and firmware)".
In the text box, type your model, ie: laserjet 2420d in your case.
Click the arrow button to the right of the text box.
At the next screen, pick your specific model if it's not already selected.
At the next screen, If a link named Cross operating system (BIOS, Firmware, Diagnostics, etc.) is there, the firmware is flashable. Click it.
If it's not, you would have to buy a new firmware dimm from HP to upgrade the firmware.
Important - At the next screen, do not click on the Download button. Instead, click the named link for the firmware for your operating system.
At the next screen, select the Readme tab. That contains all the information from all tabs.
Important - The cautious part: Look for your specific error in the Fixes list in the Readme tab information. If it's not there, I'd recommend not installing new firmware.
You can also contact HP directly for help. Just click your country here:
http://welcome.hp.com/country/us/en/wwcontact_us.h
I understand they now charge for phone support, but that chat and email support are still free.
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